The Organization and Role During Locomotion of the Proximal Musculature of the Cricket Foreleg. II. Electromyographic Activity During Stepping Patterns

A description is made of the patterns of electrical activity in the proximal muscles of the cricket foreleg during restrained locomotion and seeking movements, while the animal is held by the mesonotum, allowing the legs complete freedom of movement. 1. The initiation of the swing phase corresponds to the onset of the abductor muscle activity (Fig. 1). Its duration is matched by that of abduction-promotion and does not depend on the step frequency. Leg position is more variable at the end of the stance than at the end of the swing. 2. The promotor and abductor muscle activities are linked (Fig. 2). At least three units can be distinguished in each and the duration of their bursts is independent of the period (Fig. 3). 3. In the double depressors of the trochanter, muscles 77-lb,c (Fig. 4), one unit per muscle was identified, bursting during the swing phase. The duration of the burst is independent of the period. Some isolated potentials occasionally occur during the stance phase. 4. The overall activity in the lateral and medial remotors is coupled to the period; three main patterns can be described, depending upon the muscle bundle and the velocity of movement (Fig. 5). 5. In the coxal depressors two patterns of activity are described which depend on velocity of stepping (Fig. 6): (i) during regular and fast stepping (at frequencies greater than 2–5 Hz), the activity is coupled to that of the double depressors; (ii) during slow or irregular stepping, the activity is biphasic: an initial burst is followed after a latency correlated to the period by a second one in the second half of the stance phase. Conversely, the latency between the end of the second burst and the onset of the following abductor burst does not depend on the period. In most cases, a fast neurone (large amplitude, short phasic activation) is recruited when a slow one reaches high rates of discharge

The Organization and Role During Locomotion of the Proximal Musculature of the Cricket Foreleg : I. Anatomy and Innervation

The structure of the proximal segments of the cricket (Gryllus bimaculatus) foreleg, together with the associated musculature and its innervation are described. The morphology of 50 motor neurones involved in the control of this musculature has been revealed using backfilling techniques with cobalt, horseradish peroxidase and Lucifer Yellow. The ‘ball and socket’ pleurocoxal joint is moved by three sets of anatomical antagonists (promotor-remotor, abductor-adductor, anterior-posterior rotator muscles) inserted on each side of the three axes of rotation. The axial coxotrochanteral joint is moved by the intrinsic levator and the depressor muscles; these depressors are composed of an intrinsic (coxotrochanteral) and a ‘double’ (pleurotrochanteral) subgroup. The double depressors, and all the muscles inserting on the trochantin (promotors) or the anterior coxal rim (adductor, abductors, anterior rotators) are supplied by at least eighteen neurones, whose axons run in nerve 3. The muscles that insert on the posterior coxal rim (remotors, posterior rotators) are innervated by at least twelve similar neurones whose axons run in nerve 4. The intrinsic coxal muscles are supplied by branches of nerve 5 (ten motor neurones to the levators, two to the depressors). Three presumably common inhibitors, and one Dorsal Unpaired Median (DUM) neurone have also been found

The cost of exactly simulating quantum entanglement with classical communication

We investigate the amount of communication that must augment classical local hidden variable models in order to simulate the behaviour of entangled quantum systems. We consider the scenario where a bipartite measurement is given from a set of possibilities and the goal is to obtain exactly the same correlations that arise when the actual quantum system is measured. We show that, in the case of a single pair of qubits in a Bell state, a constant number of bits of communication is always sufficient--regardless of the number of measurements under consideration. We also show that, in the case of a system of n Bell states, a constant times 2^n bits of communication are necessary.Comment: 9 pages, LaTeX, no figure

Worker flows, job flows and establishment wage differentials : analyzing the case of France

We address the relation between establishment wage differentials and worker flows, i.e. the churning rate and the quit rate. Our analysis is based on a linked employer-employee dataset covering the French private non-farm sector from 2002 to 2005. Our estimations support the hypothesis that wage premium is an efficient human resource management tool to stabilize workers : churning rates are lower in high-paying firms due to lower quit rates. We further show that the relation is not linear, and it differs among skill groups and according to establishment size : it is strongest for low-wage levels, for low-skilled workers and in large establishments.Establishment wage effects, worker flows, churning rate, quite rate, linked employer-employee panel data, France.

Field equations and cosmology for a class of nonlocal metric models of MOND

We consider a class of nonlocal, pure-metric modified gravity models which were developed to reproduce the Tully-Fisher relation without dark matter and without changing the amount of weak lensing predicted by general relativity. Previous work gave only the weak field limiting form of the field equations specialized to a static and spherically symmetric geometry. Here we derive the full field equations and specialize them to a homogeneous, isotropic and spatially flat geometry. We also discuss the problem of fitting the free function to reproduce the expansion history. Results are derived for models in which the MOND acceleration a_0 ~ 1.2 x 10^{-10} m/s^{2} is a fundamental constant and for the more phenomenologically interesting case in which the MOND acceleration changes with the cosmological expansion rate.Comment: 15 pages, no figures, uses revtex4, dedicated to Stanley Deser on the occasion of his 83rd birthda